Ct
Deviation Score
Gene name CtxA CtxP MedO Thal MidB Stri Hipp HyTh Cebl OlfB Pitu
Ave. ΔCt
CtxA CtxP MedO Thal MidB Stri Hipp HyTh Cebl OlfB Pitu Variance
CtxA (13)


























Emx2 24.8 25.8 31.7 26.8 26.6 26.4 25.0 29.1 35.7 26.6 32.2
28.2 3.4
60.4 57.4 39.0 54.4 55.0 55.4 60.0 47.3 28.7 54.9 37.5 3.2
Neurod6 21.5 22.1 25.1 26.2 28.0 24.0 20.2 28.5 24.5 27.3 40.0
26.1 4.6
60.7 59.1 51.6 48.6 43.9 54.3 64.0 42.7 52.9 45.9 26.2 3.9
Lhx2 21.7 22.1 26.9 24.2 24.2 23.4 22.1 25.5 31.3 23.5 26.9
24.7 3.0
61.1 59.7 41.8 52.0 51.8 54.9 59.6 47.1 25.6 54.7 41.7 2.7
Gsh2 27.8 29.4 30.6 28.2 35.4 28.7 29.0 30.9 32.9 30.0 40.0
31.2 3.4
61.7 55.4 50.2 60.3 32.3 58.0 56.9 49.2 40.9 52.6 32.3 2.4
LOC218298 29.2 28.8 34.1 31.7 34.9 29.8 29.0 32.2 40.0 32.8 40.0
33.0 3.8
62.0 63.5 41.4 51.4 38.2 59.2 62.7 49.5 37.7 46.7 37.7 2.4
Egr4 20.9 21.3 26.3 23.3 25.6 21.1 22.0 25.6 23.4 22.9 27.3
23.6 2.7
62.4 60.6 37.6 51.5 40.8 61.8 57.3 40.6 51.0 53.5 32.7 2.1
Lhx6 24.5 25.2 30.1 25.9 30.1 25.3 25.3 26.3 30.2 30.2 29.9
27.5 3.0
62.7 60.1 39.0 57.0 39.2 59.5 59.4 55.3 39.0 38.9 40.0 2.4
Tbr1 19.8 20.4 28.3 24.2 29.2 21.9 21.3 24.7 27.6 23.4 31.8
24.8 5.0
63.3 61.7 40.7 51.4 38.4 57.6 59.2 50.3 42.5 53.6 31.3 3.8
Emx1 23.8 23.8 30.7 28.7 31.6 26.4 24.3 31.5 30.7 26.0 29.5
27.9 4.1
63.9 63.7 40.7 47.5 37.6 55.0 62.1 38.0 40.6 56.3 44.7 3.0
Egr3 21.3 23.0 29.2 26.0 26.9 22.7 23.8 29.3 31.6 23.7 30.5
26.2 4.9
64.5 59.3 41.2 50.6 47.8 60.4 57.1 40.6 34.0 57.4 37.2 3.4
Mef2c 19.3 20.2 24.1 21.6 23.4 21.6 22.1 24.1 22.9 20.7 25.8
22.3 3.1
66.5 61.7 40.4 53.8 44.4 54.0 51.6 40.3 46.9 58.9 31.3 1.9
Stat4 27.2 27.7 31.4 30.5 30.1 29.8 30.7 31.1 30.8 31.0 27.2
29.8 2.6
66.7 63.5 39.4 45.1 47.8 49.9 43.9 41.5 43.4 42.0 66.8 1.5
LOC226439 27.7 27.6 33.7 31.9 33.7 31.0 32.0 32.0 31.7 32.2 34.6
31.7 3.9
68.4 69.0 40.4 48.9 40.4 52.9 48.2 48.4 49.6 47.4 36.2 2.1




























CtxP (4)


























Tbr1 19.8 20.4 28.3 24.2 29.2 21.9 21.3 24.7 27.6 23.4 31.8
24.8 4.4
63.3 61.7 40.7 51.4 38.4 57.6 59.2 50.3 42.5 53.6 31.3 3.8
LOC218298 29.2 28.8 34.1 31.7 34.9 29.8 29.0 32.2 40.0 32.8 40.0
33.0 4.1
62.0 63.5 41.4 51.4 38.2 59.2 62.7 49.5 37.7 46.7 37.7 2.4
Emx1 23.8 23.8 30.7 28.7 31.6 26.4 24.3 31.5 30.7 26.0 29.5
27.9 4.1
63.9 63.7 40.7 47.5 37.6 55.0 62.1 38.0 40.6 56.3 44.7 3.0
LOC226439 27.7 27.6 33.7 31.9 33.7 31.0 32.0 32.0 31.7 32.2 34.6
31.7 4.1
68.4 69.0 40.4 48.9 40.4 52.9 48.2 48.4 49.6 47.4 36.2 2.1




























MedO (57)


























Shox2 38.4 38.8 27.9 25.5 26.3 37.9 38.3 27.3 36.1 36.8 32.9
33.3 5.4
41.8 41.8 60.0 66.1 64.1 41.8 41.8 61.6 41.8 41.8 47.3 3.9
Foxb1 32.1 34.1 26.9 27.2 25.4 32.3 35.4 24.8 29.8 40.0 40.0
31.6 4.8
46.3 41.0 60.0 59.0 63.8 45.7 38.8 65.3 52.4 38.8 38.8 3.8
Ebf2 31.3 32.8 26.3 27.8 26.1 33.4 32.2 28.0 25.0 29.1 29.1
29.2 2.9
42.2 36.8 60.6 55.2 61.1 34.7 39.1 54.2 65.2 50.5 50.2 2.8
Nkx6-2 25.3 26.2 22.9 23.6 24.3 24.7 24.8 23.7 25.3 27.1 32.0
25.4 2.5
50.4 46.9 60.6 58.0 55.0 53.2 52.8 57.3 50.6 42.8 22.3 2.4
Irx5 30.9 32.2 24.7 25.6 23.9 29.8 29.6 26.1 24.4 31.4 27.3
27.8 3.1
39.4 35.0 60.7 57.6 63.3 43.0 43.9 55.9 61.8 37.8 51.6 2.9
Lhx5 35.9 38.6 29.8 30.7 29.4 35.8 34.3 30.2 28.9 34.3 35.3
33.0 3.3
40.2 40.2 60.9 57.1 62.2 40.2 42.8 59.1 64.4 42.8 40.2 2.5
Irx3 31.8 32.2 25.8 27.0 25.7 31.2 31.6 27.3 26.1 31.5 27.8
28.9 3.1
38.9 37.2 62.0 57.3 62.5 41.2 39.5 56.3 60.7 40.0 54.4 2.6
Irx1 29.0 30.6 24.5 25.2 23.9 29.5 30.2 26.4 24.6 30.0 30.1
27.6 3.1
44.6 38.4 62.2 59.5 64.4 42.8 40.1 54.9 61.8 40.8 40.5 2.6
Onecut1 29.8 33.7 26.5 27.3 26.7 32.3 28.3 27.3 32.2 36.3 32.8
30.3 3.7
51.2 38.0 62.3 59.7 61.7 42.9 56.4 59.6 43.3 33.7 41.2 3.0
Gbx1 28.2 31.6 26.4 26.5 32.1 26.9 32.1 27.1 29.1 31.4 33.2
29.5 3.1
55.4 41.6 62.5 62.1 39.6 60.6 39.6 59.5 51.5 42.2 35.3 2.5
Pax3 40.0 40.0 27.2 29.1 27.3 40.0 40.0 32.7 25.1 39.2 40.0
34.6 7.4
41.9 41.9 62.6 57.6 62.4 41.9 41.9 48.0 68.2 41.9 41.9 3.8
En1 40.0 40.0 27.5 27.6 26.8 35.0 34.6 32.1 28.0 40.0 39.0
33.7 6.2
41.3 41.3 62.8 62.4 64.9 41.4 42.4 49.7 61.2 41.3 41.3 3.5
Nr0b2 34.6 40.0 28.0 29.9 33.1 35.3 35.3 31.5 27.0 34.2 27.9
32.5 4.5
41.1 40.0 63.0 56.7 46.2 40.0 40.0 51.4 66.2 42.5 63.1 3.0
Irx2 32.0 35.6 23.1 24.9 23.1 31.7 33.6 27.2 25.0 35.2 32.1
29.4 6.3
44.2 37.4 63.8 59.8 63.9 44.7 40.6 54.8 59.6 37.4 43.8 4.5
Esrrb 27.5 29.0 25.1 26.3 26.0 29.5 27.7 28.8 25.1 29.1 30.9
27.7 2.6
51.4 42.9 64.3 58.0 59.4 40.4 49.9 43.9 64.4 42.3 32.9 1.8
Wt1 29.5 30.1 28.3 31.8 34.3 31.6 31.0 33.9 32.5 28.7 37.0
31.7 3.4
59.2 56.8 64.8 48.8 37.1 49.6 52.5 39.0 45.4 63.1 33.8 2.1
Gata3 34.4 34.0 24.5 24.8 23.3 33.4 31.8 30.9 31.6 33.3 31.5
30.3 5.8
39.4 40.6 64.9 64.1 68.0 42.1 46.2 48.6 46.7 42.4 47.0 3.9
Pou4f2 40.0 40.0 28.2 31.8 25.3 40.0 33.4 40.0 33.8 40.0 40.0
35.7 7.5
43.5 43.5 65.1 53.7 74.4 43.5 48.5 43.5 47.4 43.5 43.5 3.2
Ebf3 32.5 33.8 26.7 28.4 27.1 33.0 32.4 29.1 28.3 30.8 33.0
30.5 3.8
42.0 36.7 65.2 58.4 63.4 39.9 42.1 55.3 58.7 48.7 39.6 2.5
Dmbx1 32.0 31.4 27.6 30.5 27.1 35.5 33.8 34.6 30.0 40.0 34.8
32.5 4.9
49.8 52.0 65.7 55.4 67.5 39.2 43.4 40.7 57.1 39.2 40.1 2.8
Nkx2-9 32.6 34.9 27.9 29.0 30.5 30.0 33.6 31.0 29.3 32.4 37.3
31.7 3.8
45.2 35.1 65.8 60.6 54.3 56.5 40.6 51.9 59.4 46.0 34.6 2.3
Hmx3* 37.8 40.0 28.7 32.4 32.4 36.8 36.2 26.7 37.2 40.0 40.0
35.3 6.6
43.6 43.6 66.0 52.9 53.0 43.6 43.6 73.1 43.6 43.6 43.6 2.8
Foxd3 32.2 40.0 27.1 35.9 33.9 40.0 34.8 35.6 33.4 24.6 33.1
33.7 6.6
51.4 43.0 66.5 43.0 46.4 43.0 43.5 43.0 47.7 73.9 48.7 3.4
Pax5 33.8 35.7 28.2 29.2 27.9 34.8 34.5 35.6 33.0 34.5 33.1
32.8 4.5
45.7 41.2 66.5 63.0 67.8 41.7 43.1 41.2 48.6 43.1 48.1 2.7
Pou4f1 33.4 34.6 26.1 26.8 26.3 34.6 34.2 33.0 33.3 40.0 33.0
32.3 6.2
45.5 42.0 67.0 64.9 66.2 41.9 43.1 46.6 45.6 40.7 46.5 3.4
Lmx1b 31.7 32.6 26.2 27.9 29.0 33.7 33.2 28.3 31.0 31.0 30.2
30.4 4.2
44.3 40.3 68.7 61.1 56.6 35.6 37.9 59.4 47.7 47.5 51.0 2.3
Pou4f3 34.0 35.5 28.7 32.8 28.2 33.8 40.0 40.0 40.0 34.4 40.0
35.2 6.5
47.3 43.2 69.4 52.2 71.4 48.2 43.2 43.2 43.2 45.7 43.2 2.4
Irx4 35.1 40.0 26.2 30.6 27.8 40.0 31.2 34.5 31.5 35.3 34.6
33.3 7.2
41.5 41.5 70.4 56.1 65.1 41.5 53.8 43.0 53.0 41.5 42.7 3.0
Olig3 33.7 31.9 29.7 31.3 34.2 35.9 34.1 32.0 32.7 36.6 35.7
33.4 3.8
46.7 57.6 70.6 60.7 43.7 39.0 44.3 56.9 52.7 39.0 39.0 1.7
Pax2 40.0 40.0 23.8 31.4 30.4 40.0 35.1 31.1 25.2 40.0 40.0
34.3 10.5
42.4 42.4 70.8 51.4 54.1 42.4 42.4 52.2 67.2 42.4 42.4 4.0
Pax8 34.4 35.6 24.8 28.0 28.0 35.2 32.2 32.5 30.5 32.7 40.0
32.2 7.4
41.5 39.8 70.9 61.1 61.1 39.8 48.5 47.3 53.5 46.8 39.8 3.3
Evx1 40.0 40.0 26.5 32.4 28.0 40.0 36.2 40.0 32.1 40.0 40.0
35.9 9.4
43.6 43.6 72.2 52.3 67.3 43.6 43.6 43.6 53.3 43.6 43.6 3.0
Hoxd1 31.9 32.6 26.5 30.2 30.3 29.9 31.5 31.9 30.4 31.1 40.0
31.5 5.0
45.4 42.0 72.9 54.2 53.5 55.5 47.7 45.8 53.4 49.6 30.1 2.0
Chx10 40.0 40.0 27.1 32.1 29.7 40.0 40.0 36.2 40.0 35.5 40.0
36.4 9.3
44.4 44.4 74.6 55.4 64.6 44.4 44.4 44.4 44.4 44.4 44.4 2.6
Phox2a 34.8 40.0 28.3 36.5 31.3 34.4 35.8 34.6 33.4 36.4 34.3
34.5 6.2
44.7 43.7 76.9 43.7 62.2 46.9 43.7 45.5 51.9 43.7 47.2 2.0
Hoxa2 40.0 40.0 24.9 33.8 33.3 40.0 40.0 33.6 29.4 40.0 40.0
35.9 11.0
44.0 44.0 77.0 48.1 49.7 44.0 44.0 48.7 62.4 44.0 44.0 3.1
Hoxa7 31.2 33.2 25.3 34.9 33.7 32.1 31.5 34.5 32.9 34.4 35.7
32.7 7.3
55.3 47.6 77.7 41.2 45.7 52.2 54.1 42.9 49.0 43.3 40.9 2.6
Hoxd3 40.0 40.0 25.8 36.0 35.6 36.4 40.0 35.9 30.3 35.4 40.0
36.0 10.2
45.6 45.6 78.0 45.6 45.6 45.6 45.6 45.6 61.9 45.6 45.6 2.9
Hoxa4 40.0 40.0 23.3 40.0 34.8 40.0 40.0 34.5 29.5 40.0 40.0
36.5 13.3
45.4 45.4 78.4 45.4 46.1 45.4 45.4 46.8 61.0 45.4 45.4 3.5
Hoxa3 40.0 40.0 24.9 37.7 40.0 40.0 40.0 40.0 30.3 40.0 40.0
37.5 12.7
45.6 45.6 78.4 45.6 45.6 45.6 45.6 45.6 61.0 45.6 45.6 3.1
Hoxb5 40.0 37.5 23.2 37.4 35.8 40.0 35.6 40.0 29.6 35.9 37.3
35.7 12.5
45.6 45.6 78.6 45.6 45.6 45.6 45.6 45.6 60.6 45.6 45.6 3.6
Phox2b 40.0 40.0 26.1 33.8 31.3 39.1 40.0 37.6 33.3 38.7 37.3
36.1 10.0
44.6 44.6 78.6 49.1 58.8 44.6 44.6 44.6 51.0 44.6 44.6 2.6
Hoxa5 36.4 40.0 24.0 40.0 35.4 37.1 36.1 39.6 30.3 40.0 37.5
36.1 12.0
45.7 45.7 78.9 45.7 45.7 45.7 45.7 45.7 59.9 45.7 45.7 3.3
Hoxb3 40.0 40.0 24.7 37.0 35.1 40.0 40.0 40.0 30.7 40.0 40.0
37.0 12.3
45.7 45.7 79.0 45.7 45.7 45.7 45.7 45.7 59.5 45.7 45.7 3.1
Hoxd8 35.4 34.7 26.8 35.0 32.7 35.6 34.3 40.0 32.5 34.5 35.7
34.3 7.5
44.4 45.4 79.3 44.4 54.3 44.4 47.4 44.4 55.1 46.7 44.4 2.3
Hoxb4 34.9 36.4 26.6 35.1 37.1 34.6 35.5 35.3 31.7 35.8 35.0
34.4 7.8
45.8 45.6 79.3 45.6 45.6 47.1 45.6 45.6 58.7 45.6 45.6 2.5
Hoxc4 40.0 40.0 25.4 40.0 37.5 40.0 40.0 40.0 32.6 40.0 32.4
37.1 11.7
45.2 45.2 79.7 45.2 45.2 45.2 45.2 45.2 53.8 45.2 54.7 2.8
Hoxc5 40.0 40.0 25.2 36.2 35.2 40.0 40.0 40.0 32.0 40.0 34.6
36.7 11.4
45.8 45.8 80.2 45.8 45.8 45.8 45.8 45.8 56.3 45.8 47.0 2.8
Hoxb8 40.0 40.0 23.0 40.0 35.7 40.0 40.0 40.0 32.1 35.1 37.4
36.7 13.6
46.1 46.1 80.7 46.1 46.1 46.1 46.1 46.1 54.6 46.1 46.1 3.5
Hoxd4 37.3 39.2 25.8 38.1 40.0 39.6 40.0 40.0 33.1 37.3 34.5
36.8 11.0
46.0 46.0 80.9 46.0 46.0 46.0 46.0 46.0 53.3 46.0 47.8 2.6
Hoxb6 40.0 40.0 25.2 40.0 40.0 40.0 40.0 40.0 33.3 40.0 40.0
38.1 12.8
46.3 46.3 81.2 46.3 46.3 46.3 46.3 46.3 52.2 46.3 46.3 2.8
Hoxa6 40.0 40.0 25.7 40.0 40.0 40.0 40.0 40.0 33.7 40.0 40.0
38.1 12.4
46.4 46.4 81.3 46.4 46.4 46.4 46.4 46.4 51.2 46.4 46.4 2.7
Hoxc8 40.0 40.0 29.0 40.0 40.0 35.2 40.0 40.0 34.5 40.0 40.0
38.1 9.0
46.6 46.6 81.5 46.6 46.6 46.6 46.6 46.6 49.4 46.6 46.6 1.7
Hoxb9 40.0 40.0 29.7 40.0 37.0 35.0 40.0 40.0 40.0 40.0 40.0
38.3 8.6
46.8 46.8 81.6 46.8 46.8 46.8 46.8 46.8 46.8 46.8 46.8 1.5
Hoxc6 40.0 40.0 27.4 40.0 39.3 40.0 40.0 37.0 35.1 40.0 40.0
38.1 10.7
46.8 46.8 81.6 46.8 46.8 46.8 46.8 46.8 46.8 46.8 46.8 2.2
Hoxb7 40.0 40.0 27.2 40.0 40.0 40.0 40.0 40.0 35.5 40.0 40.0
38.4 11.2
46.8 46.8 81.6 46.8 46.8 46.8 46.8 46.8 46.8 46.8 46.8 2.2
4933413G11Rik 40.0 40.0 25.2 40.0 39.2 39.5 40.0 40.0 35.4 37.4 40.0
37.9 12.7
46.8 46.8 81.6 46.8 46.8 46.8 46.8 46.8 46.8 46.8 46.8 2.8




























Thal (24)


























Isl1 27.6 30.9 27.2 25.4 30.2 25.8 33.1 25.1 34.9 34.1 25.5
29.1 3.7
54.3 44.7 55.2 60.3 46.9 59.2 38.5 61.2 33.6 35.9 60.1 3.5
Gsh2 27.8 29.4 30.6 28.2 35.4 28.7 29.0 30.9 32.9 30.0 40.0
31.2 3.0
61.7 55.4 50.2 60.3 32.3 58.0 56.9 49.2 40.9 52.6 32.3 2.4
Sim1 30.1 32.5 31.8 29.3 31.4 34.6 35.8 25.8 34.5 40.0 34.6
32.8 3.4
57.6 49.1 51.6 60.4 53.1 41.6 40.2 72.9 41.8 40.2 41.6 2.8
Nkx2-9 32.6 34.9 27.9 29.0 30.5 30.0 33.6 31.0 29.3 32.4 37.3
31.7 2.6
45.2 35.1 65.8 60.6 54.3 56.5 40.6 51.9 59.4 46.0 34.6 2.3
Pax8 34.4 35.6 24.8 28.0 28.0 35.2 32.2 32.5 30.5 32.7 40.0
32.2 4.2
41.5 39.8 70.9 61.1 61.1 39.8 48.5 47.3 53.5 46.8 39.8 3.3
Lmx1b 31.7 32.6 26.2 27.9 29.0 33.7 33.2 28.3 31.0 31.0 30.2
30.4 2.5
44.3 40.3 68.7 61.1 56.6 35.6 37.9 59.4 47.7 47.5 51.0 2.3
Rarb 26.8 28.9 29.3 26.3 29.0 23.9 31.6 30.5 32.5 32.1 30.2
29.2 2.9
59.5 51.1 49.6 61.3 50.9 71.0 40.6 44.7 36.9 38.5 45.9 2.5
Lhx9 28.5 28.9 26.3 23.7 22.5 31.8 24.5 25.4 28.3 28.8 33.3
27.5 3.8
46.7 45.4 53.7 61.9 65.4 36.3 59.3 56.4 47.2 46.0 31.7 3.2
Gbx1 28.2 31.6 26.4 26.5 32.1 26.9 32.1 27.1 29.1 31.4 33.2
29.5 3.0
55.4 41.6 62.5 62.1 39.6 60.6 39.6 59.5 51.5 42.2 35.3 2.5
En1 40.0 40.0 27.5 27.6 26.8 35.0 34.6 32.1 28.0 40.0 39.0
33.7 6.1
41.3 41.3 62.8 62.4 64.9 41.4 42.4 49.7 61.2 41.3 41.3 3.5
Pax5 33.8 35.7 28.2 29.2 27.9 34.8 34.5 35.6 33.0 34.5 33.1
32.8 3.6
45.7 41.2 66.5 63.0 67.8 41.7 43.1 41.2 48.6 43.1 48.1 2.7
Lhx8 26.4 30.1 33.4 24.7 35.0 24.4 33.1 25.0 35.0 34.5 30.4
30.2 5.5
59.2 50.2 42.1 63.3 38.4 63.9 42.8 62.5 38.5 39.6 49.5 4.1
Gata3 34.4 34.0 24.5 24.8 23.3 33.4 31.8 30.9 31.6 33.3 31.5
30.3 5.5
39.4 40.6 64.9 64.1 68.0 42.1 46.2 48.6 46.7 42.4 47.0 3.9
Nkx3-1 29.7 31.7 33.9 28.2 31.0 30.8 30.7 30.1 32.4 28.4 40.0
31.5 3.4
56.7 46.9 36.1 64.6 50.2 51.2 51.9 55.0 43.3 63.6 30.5 2.0
Pou4f1 33.4 34.6 26.1 26.8 26.3 34.6 34.2 33.0 33.3 40.0 33.0
32.3 5.5
45.5 42.0 67.0 64.9 66.2 41.9 43.1 46.6 45.6 40.7 46.5 3.4
Gtf2a1lf 30.4 29.8 30.5 27.4 29.5 29.9 33.3 28.5 32.4 35.0 29.4
30.5 3.1
50.9 53.8 50.4 64.9 55.0 53.0 36.8 59.6 41.3 28.7 55.6 2.1
Shox2 38.4 38.8 27.9 25.5 26.3 37.9 38.3 27.3 36.1 36.8 32.9
33.3 7.8
41.8 41.8 60.0 66.1 64.1 41.8 41.8 61.6 41.8 41.8 47.3 3.9
Gbx2 30.4 34.3 27.1 24.5 29.2 29.1 34.1 24.8 32.1 33.3 40.0
30.8 6.3
50.0 39.1 59.1 66.3 53.2 53.5 39.5 65.4 45.2 41.7 37.0 3.6
Foxa2 40.0 40.0 30.8 28.0 29.0 40.0 40.0 29.8 40.0 40.0 35.2
35.7 7.7
42.6 42.6 58.0 67.9 64.3 42.6 42.6 61.5 42.6 42.6 42.6 2.8
Pitx3 33.0 34.3 30.0 27.8 29.3 33.5 34.6 29.7 32.7 35.4 40.0
32.7 5.0
47.0 41.9 59.2 68.1 62.0 45.1 40.6 60.2 48.1 38.9 38.9 2.5
Foxa1 40.0 40.0 32.3 29.4 31.8 40.0 40.0 30.1 40.0 40.0 32.5
36.0 6.6
41.7 41.7 54.7 69.0 57.3 41.7 41.7 65.3 41.7 41.7 53.7 2.1
Tcf7l2 24.6 25.8 24.3 21.2 21.7 25.2 24.6 22.7 24.7 24.5 24.5
24.0 2.8
45.2 37.0 47.8 69.8 66.6 41.5 45.8 59.4 44.5 46.1 46.3 1.4
Lef1 24.3 25.1 24.5 21.8 23.0 24.8 24.6 23.7 24.7 24.5 27.1
24.4 2.5
50.6 44.4 49.1 70.3 61.2 46.0 48.3 55.6 47.4 48.8 28.3 1.2
Msx3 31.4 32.2 33.1 27.4 35.3 31.7 32.0 33.9 32.5 29.1 34.0
32.0 4.6
53.1 49.3 44.7 72.1 35.7 51.6 49.9 41.2 47.9 63.9 40.5 2.1




























MidB (32)


























Pitx1 34.9 36.2 33.4 32.2 28.4 36.2 35.6 35.1 36.5 33.1 21.8
33.0 4.6
44.1 44.0 48.1 51.1 60.6 44.0 44.0 44.0 44.0 48.7 77.6 3.9
Pax8 34.4 35.6 24.8 28.0 28.0 35.2 32.2 32.5 30.5 32.7 40.0
32.2 4.1
41.5 39.8 70.9 61.1 61.1 39.8 48.5 47.3 53.5 46.8 39.8 3.3
Ebf2 31.3 32.8 26.3 27.8 26.1 33.4 32.2 28.0 25.0 29.1 29.1
29.2 3.1
42.2 36.8 60.6 55.2 61.1 34.7 39.1 54.2 65.2 50.5 50.2 2.8
Onecut1 29.8 33.7 26.5 27.3 26.7 32.3 28.3 27.3 32.2 36.3 32.8
30.3 3.6
51.2 38.0 62.3 59.7 61.7 42.9 56.4 59.6 43.3 33.7 41.2 3.0
Otx2 31.1 31.0 25.7 24.4 23.7 29.7 25.7 25.8 23.3 28.2 26.8
26.8 3.1
34.2 34.2 54.3 59.3 61.9 39.2 54.3 54.1 63.3 44.9 50.2 2.7
Pitx3 33.0 34.3 30.0 27.8 29.3 33.5 34.6 29.7 32.7 35.4 40.0
32.7 3.5
47.0 41.9 59.2 68.1 62.0 45.1 40.6 60.2 48.1 38.9 38.9 2.5
Lhx5 35.9 38.6 29.8 30.7 29.4 35.8 34.3 30.2 28.9 34.3 35.3
33.0 3.6
40.2 40.2 60.9 57.1 62.2 40.2 42.8 59.1 64.4 42.8 40.2 2.5
Pax3 40.0 40.0 27.2 29.1 27.3 40.0 40.0 32.7 25.1 39.2 40.0
34.6 7.3
41.9 41.9 62.6 57.6 62.4 41.9 41.9 48.0 68.2 41.9 41.9 3.8
Irx3 31.8 32.2 25.8 27.0 25.7 31.2 31.6 27.3 26.1 31.5 27.8
28.9 3.2
38.9 37.2 62.0 57.3 62.5 41.2 39.5 56.3 60.7 40.0 54.4 2.6
Irx5 30.9 32.2 24.7 25.6 23.9 29.8 29.6 26.1 24.4 31.4 27.3
27.8 3.9
39.4 35.0 60.7 57.6 63.3 43.0 43.9 55.9 61.8 37.8 51.6 2.9
Ebf3 32.5 33.8 26.7 28.4 27.1 33.0 32.4 29.1 28.3 30.8 33.0
30.5 3.3
42.0 36.7 65.2 58.4 63.4 39.9 42.1 55.3 58.7 48.7 39.6 2.5
Foxb1 32.1 34.1 26.9 27.2 25.4 32.3 35.4 24.8 29.8 40.0 40.0
31.6 6.2
46.3 41.0 60.0 59.0 63.8 45.7 38.8 65.3 52.4 38.8 38.8 3.8
Irx2 32.0 35.6 23.1 24.9 23.1 31.7 33.6 27.2 25.0 35.2 32.1
29.4 6.3
44.2 37.4 63.8 59.8 63.9 44.7 40.6 54.8 59.6 37.4 43.8 4.5
Shox2 38.4 38.8 27.9 25.5 26.3 37.9 38.3 27.3 36.1 36.8 32.9
33.3 7.0
41.8 41.8 60.0 66.1 64.1 41.8 41.8 61.6 41.8 41.8 47.3 3.9
Foxa2 40.0 40.0 30.8 28.0 29.0 40.0 40.0 29.8 40.0 40.0 35.2
35.7 6.7
42.6 42.6 58.0 67.9 64.3 42.6 42.6 61.5 42.6 42.6 42.6 2.8
Irx1 29.0 30.6 24.5 25.2 23.9 29.5 30.2 26.4 24.6 30.0 30.1
27.6 3.7
44.6 38.4 62.2 59.5 64.4 42.8 40.1 54.9 61.8 40.8 40.5 2.6
Chx10 40.0 40.0 27.1 32.1 29.7 40.0 40.0 36.2 40.0 35.5 40.0
36.4 6.7
44.4 44.4 74.6 55.4 64.6 44.4 44.4 44.4 44.4 44.4 44.4 2.6
Tcfap2b 27.2 28.9 25.1 27.1 23.1 31.5 31.1 29.8 22.3 26.5 28.0
27.3 4.2
50.3 44.5 57.8 50.8 64.8 35.3 36.9 41.4 67.7 53.0 47.6 2.9
En1 40.0 40.0 27.5 27.6 26.8 35.0 34.6 32.1 28.0 40.0 39.0
33.7 6.9
41.3 41.3 62.8 62.4 64.9 41.4 42.4 49.7 61.2 41.3 41.3 3.5
Pax7 33.6 34.2 26.4 28.0 24.0 40.0 40.0 29.1 32.7 34.9 23.1
31.4 7.5
43.1 41.5 59.5 55.8 65.0 39.8 39.8 53.3 45.1 40.1 67.0 4.4
Irx4 35.1 40.0 26.2 30.6 27.8 40.0 31.2 34.5 31.5 35.3 34.6
33.3 5.5
41.5 41.5 70.4 56.1 65.1 41.5 53.8 43.0 53.0 41.5 42.7 3.0
Lhx9 28.5 28.9 26.3 23.7 22.5 31.8 24.5 25.4 28.3 28.8 33.3
27.5 4.9
46.7 45.4 53.7 61.9 65.4 36.3 59.3 56.4 47.2 46.0 31.7 3.2
Pou4f1 33.4 34.6 26.1 26.8 26.3 34.6 34.2 33.0 33.3 40.0 33.0
32.3 6.0
45.5 42.0 67.0 64.9 66.2 41.9 43.1 46.6 45.6 40.7 46.5 3.4
Evx1 40.0 40.0 26.5 32.4 28.0 40.0 36.2 40.0 32.1 40.0 40.0
35.9 8.0
43.6 43.6 72.2 52.3 67.3 43.6 43.6 43.6 53.3 43.6 43.6 3.0
Dmbx1 32.0 31.4 27.6 30.5 27.1 35.5 33.8 34.6 30.0 40.0 34.8
32.5 5.4
49.8 52.0 65.7 55.4 67.5 39.2 43.4 40.7 57.1 39.2 40.1 2.8
Pax5 33.8 35.7 28.2 29.2 27.9 34.8 34.5 35.6 33.0 34.5 33.1
32.8 4.9
45.7 41.2 66.5 63.0 67.8 41.7 43.1 41.2 48.6 43.1 48.1 2.7
Gata3 34.4 34.0 24.5 24.8 23.3 33.4 31.8 30.9 31.6 33.3 31.5
30.3 7.0
39.4 40.6 64.9 64.1 68.0 42.1 46.2 48.6 46.7 42.4 47.0 3.9
Barhl1 38.4 39.0 31.8 34.9 25.4 33.4 39.9 29.1 25.6 32.3 32.0
32.9 7.5
40.6 40.6 50.0 40.9 68.6 45.3 40.6 57.7 67.9 48.4 49.4 3.4
Sox14 28.0 28.9 26.6 26.5 23.9 29.4 27.9 26.4 28.4 28.6 31.4
27.8 3.9
48.9 44.2 56.4 57.2 71.0 41.7 49.8 57.6 47.0 45.6 30.6 1.9
Pou4f3 34.0 35.5 28.7 32.8 28.2 33.8 40.0 40.0 40.0 34.4 40.0
35.2 7.0
47.3 43.2 69.4 52.2 71.4 48.2 43.2 43.2 43.2 45.7 43.2 2.4
Tal1 28.7 28.9 26.4 26.9 25.1 29.3 28.7 29.2 29.5 29.6 30.1
28.4 3.3
47.9 46.4 63.4 60.1 72.0 44.2 47.9 44.8 42.9 41.9 38.5 1.5
Pou4f2 40.0 40.0 28.2 31.8 25.3 40.0 33.4 40.0 33.8 40.0 40.0
35.7 10.4
43.5 43.5 65.1 53.7 74.4 43.5 48.5 43.5 47.4 43.5 43.5 3.2




























Stri (15)


























Six3 25.1 28.0 29.7 22.7 23.6 22.2 28.1 22.7 32.6 32.4 21.5
26.2 4.0
52.8 45.6 41.2 59.1 56.8 60.2 45.1 59.0 33.6 34.4 62.0 3.9
Dlx5 26.0 27.3 35.5 25.3 35.2 23.9 27.0 25.4 34.0 22.7 29.8
28.4 4.4
55.4 52.4 34.4 57.0 34.4 60.3 53.1 56.8 36.6 63.2 46.4 4.3
Egr3 21.3 23.0 29.2 26.0 26.9 22.7 23.8 29.3 31.6 23.7 30.5
26.2 3.5
64.5 59.3 41.2 50.6 47.8 60.4 57.1 40.6 34.0 57.4 37.2 3.4
Gbx1 28.2 31.6 26.4 26.5 32.1 26.9 32.1 27.1 29.1 31.4 33.2
29.5 2.6
55.4 41.6 62.5 62.1 39.6 60.6 39.6 59.5 51.5 42.2 35.3 2.5
Titf1 28.9 31.0 36.4 26.8 34.2 26.4 30.0 24.5 40.0 35.4 27.2
31.0 4.6
53.9 48.3 37.7 59.5 39.7 60.6 51.0 65.5 37.7 37.7 58.6 3.8
Isl1 26.6 29.8 26.7 24.3 29.0 23.9 31.2 23.7 34.9 32.7 24.5
27.9 4.0
53.8 44.8 53.4 59.8 47.1 61.0 41.1 61.6 31.2 37.1 59.3 3.7
Egr4 20.9 21.3 26.3 23.3 25.6 21.1 22.0 25.6 23.4 22.9 27.3
23.6 2.5
62.4 60.6 37.6 51.5 40.8 61.8 57.3 40.6 51.0 53.5 32.7 2.1
Egr2 24.6 24.2 29.3 27.2 28.9 24.1 26.4 29.3 30.0 24.5 26.4
26.8 2.7
60.2 61.9 38.4 48.5 40.2 62.4 51.9 38.4 35.4 60.6 52.1 2.2
Tcfl5 27.5 27.0 30.1 27.6 31.8 26.5 29.1 30.3 28.8 30.1 33.9
29.3 2.8
58.7 61.2 46.6 58.3 38.5 63.1 51.1 45.5 52.5 46.3 28.3 2.1
Lhx8 26.4 30.1 33.4 24.7 35.0 24.4 33.1 25.0 35.0 34.5 30.4
30.2 5.8
59.2 50.2 42.1 63.3 38.4 63.9 42.8 62.5 38.5 39.6 49.5 4.1
Mrg1 21.6 23.0 21.8 22.5 22.1 20.1 23.9 24.9 25.6 19.4 24.2
22.6 2.6
55.6 48.2 54.3 50.8 53.0 64.1 43.2 37.9 33.7 67.6 41.5 1.8
Dlx6 34.3 36.0 40.0 32.1 40.0 28.7 34.5 29.9 40.0 27.5 37.2
34.6 5.8
45.0 42.4 42.4 52.9 42.4 65.2 44.2 60.9 42.4 69.8 42.4 2.7
Rarb 26.8 28.9 29.3 26.3 29.0 23.9 31.6 30.5 32.5 32.1 30.2
29.2 5.3
59.5 51.1 49.6 61.3 50.9 71.0 40.6 44.7 36.9 38.5 45.9 2.5
Rxrg 25.1 26.0 26.2 24.9 26.0 21.4 26.1 24.7 29.0 25.6 23.2
25.3 3.9
51.4 45.9 44.8 52.2 46.3 71.4 45.6 53.3 29.6 48.2 61.3 1.8
Foxp1 21.7 23.1 23.9 23.1 23.8 20.4 22.7 24.0 23.2 24.1 23.0
23.0 2.6
62.1 49.1 41.4 49.1 42.0 74.7 52.9 40.4 47.8 40.1 50.3 1.1




























Hipp (10)


























Emx2 24.8 25.8 31.7 26.8 26.6 26.4 25.0 29.1 35.7 26.6 32.2
28.2 3.3
60.4 57.4 39.0 54.4 55.0 55.4 60.0 47.3 28.7 54.9 37.5 3.2
Foxg1 20.4 22.2 32.3 22.8 32.3 20.5 20.2 24.2 32.8 21.2 28.9
25.3 5.1
59.8 56.1 35.9 54.9 35.8 59.6 60.2 52.1 34.9 58.1 42.6 5.0
Neurod2 21.7 22.3 24.2 26.2 28.5 24.9 21.1 27.7 20.7 26.3 35.0
25.3 4.3
59.0 57.6 52.8 47.9 42.1 50.9 60.7 43.9 61.7 47.6 25.7 4.0
Og9x 25.7 27.3 27.3 28.7 30.0 28.0 25.0 29.8 24.3 29.9 35.4
28.3 3.3
59.0 53.4 53.5 48.5 43.8 51.1 61.6 44.6 64.0 44.3 26.3 2.8
Emx1 23.8 23.8 30.7 28.7 31.6 26.4 24.3 31.5 30.7 26.0 29.5
27.9 3.6
63.9 63.7 40.7 47.5 37.6 55.0 62.1 38.0 40.6 56.3 44.7 3.0
Zfp238 20.4 20.9 22.2 22.7 26.9 22.1 19.5 23.9 19.3 23.4 25.4
22.4 2.9
59.1 56.6 51.0 49.0 30.4 51.6 62.7 43.5 63.7 45.5 36.8 2.3
LOC218298 29.2 28.8 34.1 31.7 34.9 29.8 29.0 32.2 40.0 32.8 40.0
33.0 4.0
62.0 63.5 41.4 51.4 38.2 59.2 62.7 49.5 37.7 46.7 37.7 2.4
Neurod6 21.5 22.1 25.1 26.2 28.0 24.0 20.2 28.5 24.5 27.3 40.0
26.1 5.9
60.7 59.1 51.6 48.6 43.9 54.3 64.0 42.7 52.9 45.9 26.2 3.9
Nhlh1 29.1 30.2 30.2 30.7 32.2 30.7 28.0 32.0 32.6 33.3 33.5
31.1 3.1
62.2 55.9 55.6 52.5 43.5 52.4 68.9 45.0 41.0 37.0 35.9 1.7
Nr3c2 24.3 24.2 24.1 24.1 24.4 24.2 21.6 25.7 24.4 24.2 24.0
24.1 2.5
47.6 48.6 50.0 49.9 46.5 49.2 77.6 32.4 47.3 49.4 51.4 0.9




























HyTh (22)


























Pitx3 33.0 34.3 30.0 27.8 29.3 33.5 34.6 29.7 32.7 35.4 40.0
32.7 3.0
47.0 41.9 59.2 68.1 62.0 45.1 40.6 60.2 48.1 38.9 38.9 2.5
Tbx19 35.2 33.7 35.3 35.6 40.0 40.0 34.8 29.4 40.0 34.4 22.7
34.6 5.2
45.0 48.6 45.0 45.0 45.0 45.0 45.7 60.2 45.0 46.6 78.6 3.7
Dlx6 34.3 36.0 40.0 32.1 40.0 28.7 34.5 29.9 40.0 27.5 37.2
34.6 4.7
45.0 42.4 42.4 52.9 42.4 65.2 44.2 60.9 42.4 69.8 42.4 2.7
Isl1 27.6 30.9 27.2 25.4 30.2 25.8 33.1 25.1 34.9 34.1 25.5
29.1 4.0
54.3 44.7 55.2 60.3 46.9 59.2 38.5 61.2 33.6 35.9 60.1 3.5
Foxa2 40.0 40.0 30.8 28.0 29.0 40.0 40.0 29.8 40.0 40.0 35.2
35.7 5.9
42.6 42.6 58.0 67.9 64.3 42.6 42.6 61.5 42.6 42.6 42.6 2.8
Pitx2 40.0 40.0 28.7 25.1 25.2 34.5 40.0 24.1 34.5 32.5 20.8
31.4 7.3
40.3 40.3 52.6 59.7 59.5 41.4 40.3 61.6 41.2 45.2 67.9 5.1
Vax1 28.5 29.2 32.7 27.7 32.2 28.5 28.5 26.2 34.0 27.2 26.0
29.2 2.9
52.6 49.9 36.0 55.8 38.0 52.7 52.5 61.6 31.0 57.7 62.2 2.5
Shox2 38.4 38.8 27.9 25.5 26.3 37.9 38.3 27.3 36.1 36.8 32.9
33.3 6.0
41.8 41.8 60.0 66.1 64.1 41.8 41.8 61.6 41.8 41.8 47.3 3.9
Lhx8 26.4 30.1 33.4 24.7 35.0 24.4 33.1 25.0 35.0 34.5 30.4
30.2 5.2
59.2 50.2 42.1 63.3 38.4 63.9 42.8 62.5 38.5 39.6 49.5 4.1
Nr0b1 33.4 33.8 31.2 30.6 30.6 35.3 34.7 28.0 33.8 40.0 25.7
32.5 4.5
45.2 43.7 52.6 54.7 54.8 39.7 40.6 63.7 43.9 39.7 71.4 2.9
Six6 40.0 40.0 35.0 40.0 40.0 40.0 33.0 25.4 40.0 31.8 20.1
35.0 9.6
44.3 44.3 44.3 44.3 44.3 44.3 48.6 64.5 44.3 51.0 75.7 4.7
Foxb1 32.1 34.1 26.9 27.2 25.4 32.3 35.4 24.8 29.8 40.0 40.0
31.6 6.8
46.3 41.0 60.0 59.0 63.8 45.7 38.8 65.3 52.4 38.8 38.8 3.8
Gbx2 30.4 34.3 27.1 24.5 29.2 29.1 34.1 24.8 32.1 33.3 40.0
30.8 6.0
50.0 39.1 59.1 66.3 53.2 53.5 39.5 65.4 45.2 41.7 37.0 3.6
Titf1 28.9 31.0 36.4 26.8 34.2 26.4 30.0 24.5 40.0 35.4 27.2
31.0 6.4
53.9 48.3 37.7 59.5 39.7 60.6 51.0 65.5 37.7 37.7 58.6 3.8
Lmx1a 40.0 38.2 30.9 30.6 31.7 34.7 32.1 28.2 28.3 35.6 33.5
33.1 4.9
39.0 39.0 55.7 56.6 52.5 40.1 50.8 66.5 65.9 39.0 44.9 2.5
Rax 33.2 34.7 40.0 32.5 35.2 37.2 34.2 27.2 37.1 34.3 24.9
33.7 6.5
48.9 44.4 43.4 51.0 43.4 43.4 45.7 67.0 43.4 45.5 73.8 3.3
Csrp3 33.1 34.2 32.1 32.3 32.2 40.0 33.3 29.4 29.1 32.9 33.8
33.0 3.5
46.6 40.2 52.0 51.4 51.5 35.5 45.2 67.8 69.4 47.6 42.7 1.7
Sim1 30.1 32.5 31.8 29.3 31.4 34.6 35.8 25.8 34.5 40.0 34.6
32.8 6.9
57.6 49.1 51.6 60.4 53.1 41.6 40.2 72.9 41.8 40.2 41.6 2.8
Hmx3 37.8 40.0 28.7 32.4 32.4 36.8 36.2 26.7 37.2 40.0 40.0
35.3 8.6
43.6 43.6 66.0 52.9 53.0 43.6 43.6 73.1 43.6 43.6 43.6 2.8
Gcm2 40.0 40.0 40.0 31.1 40.0 40.0 40.0 29.5 40.0 40.0 40.0
38.2 8.8
45.4 45.4 45.4 66.5 45.4 45.4 45.4 75.2 45.4 45.4 45.4 1.9
Foxb2 36.9 40.0 30.8 34.9 34.5 40.0 40.0 28.5 35.9 40.0 37.2
36.3 7.7
45.1 45.1 65.3 45.5 47.4 45.1 45.1 76.0 45.1 45.1 45.1 2.1
Bsx-pending 34.9 36.3 36.3 32.0 37.6 38.8 36.3 28.1 40.0 35.9 35.7
35.6 7.5
46.3 45.6 45.6 60.0 45.6 45.6 45.6 78.8 45.6 45.6 45.6 2.1




























Cebl (49)


























Hoxb5 40.0 37.5 23.2 37.4 35.8 40.0 35.6 40.0 29.6 35.9 37.3
35.7 6.0
45.6 45.6 78.6 45.6 45.6 45.6 45.6 45.6 60.6 45.6 45.6 3.6
Irx3 31.8 32.2 25.8 27.0 25.7 31.2 31.6 27.3 26.1 31.5 27.8
28.9 2.8
38.9 37.2 62.0 57.3 62.5 41.2 39.5 56.3 60.7 40.0 54.4 2.6
Dlx3 32.6 35.9 31.9 31.0 30.2 36.0 34.2 32.9 30.0 33.8 28.2
32.4 2.5
48.5 37.0 51.7 55.8 59.6 37.0 40.9 46.9 60.8 42.6 69.2 2.1
Hoxa4 40.0 40.0 23.3 40.0 34.8 40.0 40.0 34.5 29.5 40.0 40.0
36.5 7.1
45.4 45.4 78.4 45.4 46.1 45.4 45.4 46.8 61.0 45.4 45.4 3.5
En1 40.0 40.0 27.5 27.6 26.8 35.0 34.6 32.1 28.0 40.0 39.0
33.7 5.6
41.3 41.3 62.8 62.4 64.9 41.4 42.4 49.7 61.2 41.3 41.3 3.5
Neurod2 21.7 22.3 24.2 26.2 28.5 24.9 21.1 27.7 20.7 26.3 35.0
25.3 4.6
59.0 57.6 52.8 47.9 42.1 50.9 60.7 43.9 61.7 47.6 25.7 4.0
Irx5 30.9 32.2 24.7 25.6 23.9 29.8 29.6 26.1 24.4 31.4 27.3
27.8 3.5
39.4 35.0 60.7 57.6 63.3 43.0 43.9 55.9 61.8 37.8 51.6 2.9
Irx1 29.0 30.6 24.5 25.2 23.9 29.5 30.2 26.4 24.6 30.0 30.1
27.6 3.0
44.6 38.4 62.2 59.5 64.4 42.8 40.1 54.9 61.8 40.8 40.5 2.6
Hoxa2 40.0 40.0 24.9 33.8 33.3 40.0 40.0 33.6 29.4 40.0 40.0
35.9 6.5
44.0 44.0 77.0 48.1 49.7 44.0 44.0 48.7 62.4 44.0 44.0 3.1
Madh1 27.3 29.6 27.1 27.0 23.3 23.6 27.8 23.5 22.2 23.4 23.0
25.2 3.0
41.6 31.9 42.1 42.7 58.2 57.0 39.4 57.3 62.5 57.9 59.4 2.4
Snai1 32.4 35.0 32.6 32.5 32.9 30.6 34.2 30.9 29.1 29.2 28.9
31.7 2.6
46.1 33.2 45.1 45.8 44.0 55.4 37.2 53.9 63.0 62.3 64.0 2.0
Otx2 31.1 31.0 25.7 24.4 23.7 29.7 25.7 25.8 23.3 28.2 26.8
26.8 3.5
34.2 34.2 54.3 59.3 61.9 39.2 54.3 54.1 63.3 44.9 50.2 2.7
Zfp238 20.4 20.9 22.2 22.7 26.9 22.1 19.5 23.9 19.3 23.4 25.4
22.4 3.1
59.1 56.6 51.0 49.0 30.4 51.6 62.7 43.5 63.7 45.5 36.8 2.3
Og9x 25.7 27.3 27.3 28.7 30.0 28.0 25.0 29.8 24.3 29.9 35.4
28.3 4.0
59.0 53.4 53.5 48.5 43.8 51.1 61.6 44.6 64.0 44.3 26.3 2.8
Esrrb 27.5 29.0 25.1 26.3 26.0 29.5 27.7 28.8 25.1 29.1 30.9
27.7 2.6
51.4 42.9 64.3 58.0 59.4 40.4 49.9 43.9 64.4 42.3 32.9 1.8
Lhx5 35.9 38.6 29.8 30.7 29.4 35.8 34.3 30.2 28.9 34.3 35.3
33.0 4.2
40.2 40.2 60.9 57.1 62.2 40.2 42.8 59.1 64.4 42.8 40.2 2.5
Ebf2 31.3 32.8 26.3 27.8 26.1 33.4 32.2 28.0 25.0 29.1 29.1
29.2 4.2
42.2 36.8 60.6 55.2 61.1 34.7 39.1 54.2 65.2 50.5 50.2 2.8
Lmx1a 40.0 38.2 30.9 30.6 31.7 34.7 32.1 28.2 28.3 35.6 33.5
33.1 4.7
39.0 39.0 55.7 56.6 52.5 40.1 50.8 66.5 65.9 39.0 44.9 2.5
Nhlh2 29.7 30.8 25.1 25.0 26.1 30.9 27.0 25.3 23.3 28.4 28.0
27.2 3.9
39.9 35.2 58.8 59.3 54.8 34.8 51.0 58.2 66.1 45.1 46.7 2.4
Nr0b2 34.6 40.0 28.0 29.9 33.1 35.3 35.3 31.5 27.0 34.2 27.9
32.5 5.4
41.1 40.0 63.0 56.7 46.2 40.0 40.0 51.4 66.2 42.5 63.1 3.0
Eomes 30.0 33.5 28.2 29.0 34.4 33.2 29.9 29.3 24.0 23.5 34.1
29.9 5.9
49.8 40.0 54.9 52.6 37.5 40.7 50.0 51.8 66.4 67.8 38.4 3.6
Lhx1 31.7 32.4 24.2 25.4 24.5 32.6 29.0 24.8 21.2 28.3 34.6
28.1 6.8
41.3 39.5 59.3 56.6 58.6 39.0 47.7 57.9 66.6 49.4 34.2 4.1
Meis1 26.2 27.2 23.1 25.3 24.9 24.8 27.9 25.8 22.4 23.0 25.8
25.1 2.8
43.7 37.2 62.0 49.1 51.2 52.0 33.4 46.0 66.7 62.7 46.0 1.7
Pax2 40.0 40.0 23.8 31.4 30.4 40.0 35.1 31.1 25.2 40.0 40.0
34.3 9.1
42.4 42.4 70.8 51.4 54.1 42.4 42.4 52.2 67.2 42.4 42.4 4.0
Rorc 27.9 28.3 25.3 26.3 26.8 27.4 27.2 27.2 24.1 28.9 24.3
26.7 2.6
41.7 39.3 59.5 52.7 49.1 45.0 46.7 46.7 67.4 35.6 66.2 1.5
Tcfap2b 27.2 28.9 25.1 27.1 23.1 31.5 31.1 29.8 22.3 26.5 28.0
27.3 5.0
50.3 44.5 57.8 50.8 64.8 35.3 36.9 41.4 67.7 53.0 47.6 2.9
Barhl1 38.4 39.0 31.8 34.9 25.4 33.4 39.9 29.1 25.6 32.3 32.0
32.9 7.3
40.6 40.6 50.0 40.9 68.6 45.3 40.6 57.7 67.9 48.4 49.4 3.4
E130309B19 28.7 29.6 27.4 27.6 25.5 32.3 31.3 26.6 21.8 24.7 40.0
28.7 6.9
48.7 46.3 52.3 51.8 57.6 38.4 41.3 54.6 68.1 59.9 31.0 3.6
Pax3 40.0 40.0 27.2 29.1 27.3 40.0 40.0 32.7 25.1 39.2 40.0
34.6 9.5
41.9 41.9 62.6 57.6 62.4 41.9 41.9 48.0 68.2 41.9 41.9 3.8
Trp73 30.0 30.7 30.0 27.5 30.0 29.6 27.3 28.5 26.4 28.2 30.4
28.9 2.6
42.5 37.7 42.2 60.5 42.4 45.5 62.1 53.5 68.5 55.6 39.5 1.4
Uncx4.1 31.5 33.1 26.1 27.5 26.1 32.8 33.2 26.2 22.8 31.5 33.1
29.5 6.7
44.2 39.9 59.4 55.4 59.3 40.8 39.5 59.0 68.6 44.3 39.7 3.6
Zic1 25.4 26.4 23.4 22.0 25.0 26.1 25.0 22.6 19.5 22.9 29.3
24.3 4.8
45.8 41.8 53.8 59.2 47.5 42.9 47.2 56.9 69.2 55.7 30.0 2.5
Gli 26.1 27.2 25.3 25.4 24.5 27.3 27.2 24.4 23.0 27.5 25.0
25.7 2.7
47.1 39.7 52.8 52.0 58.7 38.5 39.5 59.8 69.4 37.4 55.2 1.4
Csrp3 33.1 34.2 32.1 32.3 32.2 40.0 33.3 29.4 29.1 32.9 33.8
33.0 3.8
46.6 40.2 52.0 51.4 51.5 35.5 45.2 67.8 69.4 47.6 42.7 1.7
Dlx4 30.8 33.0 32.4 30.2 30.2 30.1 32.2 31.4 27.9 35.4 33.0
31.5 3.6
53.9 41.6 44.7 57.2 56.9 57.2 46.0 50.4 69.5 30.8 41.7 1.8
Ebf1 26.2 26.8 23.0 24.7 24.2 24.7 26.8 25.8 21.9 25.8 27.7
25.2 3.3
44.2 40.5 63.4 53.3 56.6 53.2 40.8 46.4 70.0 46.5 35.1 1.7
Zic2 25.6 26.1 24.7 24.2 25.4 27.3 25.4 25.2 20.9 24.2 30.5
25.4 4.5
49.1 47.0 53.2 55.5 50.1 41.3 49.9 51.2 70.3 55.4 27.0 2.2
Ctbp2 23.6 24.4 22.5 23.2 22.4 24.0 23.3 22.2 20.1 21.3 22.4
22.7 2.6
41.7 35.3 51.6 45.8 52.4 38.7 45.0 53.9 72.1 61.3 52.2 1.2
En2 29.7 30.5 26.0 26.6 24.9 28.8 29.1 28.8 21.0 30.8 33.7
28.2 7.2
45.3 42.8 56.6 54.9 60.0 48.2 47.1 48.1 72.1 41.9 33.0 3.3
Tcfap2a 29.1 29.9 26.5 29.3 27.1 30.6 29.8 30.4 24.2 28.7 30.9
28.8 4.6
48.2 44.4 61.7 47.1 58.6 40.9 44.8 41.8 73.4 50.1 39.0 2.0
Pax6 25.1 25.6 24.6 24.6 26.7 25.4 25.2 25.4 21.1 22.5 25.5
24.7 3.6
47.5 44.1 50.5 50.8 36.7 45.3 46.6 45.5 73.9 64.9 44.4 1.5
Bcl6 24.0 24.7 24.1 24.7 24.1 24.8 23.5 25.0 21.4 23.6 26.4
24.2 2.8
51.7 46.0 51.0 45.7 50.7 45.0 55.9 43.2 74.0 55.4 31.3 1.2
St18 25.5 26.0 23.3 23.7 24.7 24.7 23.1 25.1 20.5 26.4 24.2
24.3 3.8
42.6 39.2 56.0 53.5 47.2 47.4 57.7 44.7 74.4 36.5 50.6 1.6
Myt1 25.6 25.8 25.4 25.0 25.3 25.3 25.0 25.1 22.2 24.2 23.4
24.8 2.6
41.8 40.0 43.6 48.2 44.7 44.5 47.8 46.4 74.8 55.1 63.1 1.0
Neurod1 24.9 24.7 24.8 26.0 27.5 29.7 23.5 27.0 18.7 25.6 26.1
25.3 6.6
51.7 52.5 52.0 47.5 41.6 33.3 56.7 43.7 75.2 48.8 47.1 2.6
Max 27.8 26.7 27.6 26.9 28.6 26.0 28.6 26.5 23.3 26.6 27.5
26.9 3.7
43.7 51.3 45.0 50.1 38.3 56.5 38.3 52.7 76.2 52.5 45.5 1.4
Mad3 35.1 34.2 37.1 35.9 36.7 31.9 35.8 34.0 28.3 32.5 32.9
34.0 5.8
42.6 46.5 42.6 42.6 42.6 58.3 42.6 47.5 76.4 55.4 52.9 2.0
Hes3 29.3 29.8 27.2 30.1 30.4 30.0 33.3 29.5 20.9 31.2 31.6
29.4 8.5
50.2 48.6 57.3 47.6 46.7 48.1 37.3 49.6 77.8 44.1 42.7 3.0
Stat2 23.6 24.2 23.8 24.1 24.1 24.2 23.5 24.5 21.2 23.7 24.2
23.8 2.5
51.3 45.3 49.3 45.5 45.5 44.4 52.7 41.2 79.9 50.1 44.9 0.8




























OlfB (18)


























Hes5 28.4 32.2 29.9 28.5 29.1 27.6 29.4 27.7 31.3 26.8 39.6
30.0 3.2
55.4 38.8 49.0 54.9 52.5 58.7 51.0 58.5 42.5 62.3 26.4 2.3
Wt1 29.5 30.1 28.3 31.8 34.3 31.6 31.0 33.9 32.5 28.7 37.0
31.7 3.0
59.2 56.8 64.8 48.8 37.1 49.6 52.5 39.0 45.4 63.1 33.8 2.1
Dlx5 26.0 27.3 35.5 25.3 35.2 23.9 27.0 25.4 34.0 22.7 29.8
28.4 5.7
55.4 52.4 34.4 57.0 34.4 60.3 53.1 56.8 36.6 63.2 46.4 4.3
Nkx3-1 29.7 31.7 33.9 28.2 31.0 30.8 30.7 30.1 32.4 28.4 40.0
31.5 3.2
56.7 46.9 36.1 64.6 50.2 51.2 51.9 55.0 43.3 63.6 30.5 2.0
Msx3 31.4 32.2 33.1 27.4 35.3 31.7 32.0 33.9 32.5 29.1 34.0
32.0 2.9
53.1 49.3 44.7 72.1 35.7 51.6 49.9 41.2 47.9 63.9 40.5 2.1
Dlx1 22.8 23.6 31.3 23.4 32.3 24.1 23.1 23.2 31.3 19.9 32.1
26.1 6.1
57.5 55.8 38.2 56.2 35.8 54.5 56.8 56.6 38.2 64.0 36.4 4.4
Dlx2 26.5 27.6 34.3 26.6 40.0 26.8 27.3 26.4 35.6 23.5 40.0
30.4 6.9
57.0 54.6 38.4 56.8 36.7 56.3 55.1 57.3 36.7 64.3 36.7 4.2
Vdr 29.7 30.0 27.1 29.3 30.7 30.1 31.3 29.5 30.2 25.8 24.8
28.9 3.1
46.1 44.7 59.4 48.1 41.4 44.4 38.2 47.3 43.9 65.7 70.8 2.0
Mrg1 21.6 23.0 21.8 22.5 22.1 20.1 23.9 24.9 25.6 19.4 24.2
22.6 3.2
55.6 48.2 54.3 50.8 53.0 64.1 43.2 37.9 33.7 67.6 41.5 1.8
Eomes 30.0 33.5 28.2 29.0 34.4 33.2 29.9 29.3 24.0 23.5 34.1
29.9 6.4
49.8 40.0 54.9 52.6 37.5 40.7 50.0 51.8 66.4 67.8 38.4 3.6
Apc 25.9 29.9 26.1 26.2 27.0 25.0 26.5 25.1 23.9 22.6 27.5
26.0 3.3
50.6 28.3 49.1 48.9 44.3 55.2 47.3 54.8 61.5 68.3 41.7 1.8
Pbx3 26.0 27.0 24.3 24.4 22.9 22.3 26.9 24.1 22.9 20.5 24.5
24.2 3.6
40.1 35.0 49.0 48.8 56.9 60.0 35.6 50.2 56.9 69.2 48.3 1.9
Dlx6 34.3 36.0 40.0 32.1 40.0 28.7 34.5 29.9 40.0 27.5 37.2
34.6 7.1
45.0 42.4 42.4 52.9 42.4 65.2 44.2 60.9 42.4 69.8 42.4 2.7
Alx3 29.0 29.2 35.0 30.9 30.2 29.7 31.5 31.0 29.9 24.6 28.1
29.9 5.3
53.7 53.0 28.8 46.0 49.0 50.8 43.6 45.5 49.9 72.3 57.4 2.4
Foxd3 32.2 40.0 27.1 35.9 33.9 40.0 34.8 35.6 33.4 24.6 33.1
33.7 9.0
51.4 43.0 66.5 43.0 46.4 43.0 43.5 43.0 47.7 73.9 48.7 3.4
Alx4 32.1 34.2 34.9 33.9 32.9 36.0 33.2 37.9 32.0 29.4 40.0
34.2 4.8
57.6 45.5 41.1 46.9 53.4 40.5 51.1 40.5 58.8 73.9 40.5 1.7
Prdm1 28.0 29.2 28.9 28.9 29.4 28.8 29.4 29.6 28.6 24.9 26.7
28.4 3.5
53.2 43.9 45.9 46.5 42.5 47.2 42.8 40.9 48.6 75.8 62.5 1.3
Tcfap2c 31.1 32.5 33.7 34.9 34.9 32.7 32.0 40.0 40.0 26.7 36.4
34.1 7.4
58.0 52.2 47.1 42.3 42.4 51.6 54.4 41.9 41.9 76.2 41.9 2.4




























Pitu (39)


























Isl1 27.6 30.9 27.2 25.4 30.2 25.8 33.1 25.1 34.9 34.1 25.5
29.1 3.6
54.3 44.7 55.2 60.3 46.9 59.2 38.5 61.2 33.6 35.9 60.1 3.5
Six3 25.1 28.0 29.7 22.7 23.6 22.2 28.1 22.7 32.6 32.4 21.5
26.2 4.7
52.8 45.6 41.2 59.1 56.8 60.2 45.1 59.0 33.6 34.4 62.0 3.9
Vax1 28.5 29.2 32.7 27.7 32.2 28.5 28.5 26.2 34.0 27.2 26.0
29.2 3.1
52.6 49.9 36.0 55.8 38.0 52.7 52.5 61.6 31.0 57.7 62.2 2.5
Nr0b2 34.6 40.0 28.0 29.9 33.1 35.3 35.3 31.5 27.0 34.2 27.9
32.5 4.5
41.1 40.0 63.0 56.7 46.2 40.0 40.0 51.4 66.2 42.5 63.1 3.0
Snai1 32.4 35.0 32.6 32.5 32.9 30.6 34.2 30.9 29.1 29.2 28.9
31.7 2.8
46.1 33.2 45.1 45.8 44.0 55.4 37.2 53.9 63.0 62.3 64.0 2.0
Stat4 27.2 27.7 31.4 30.5 30.1 29.8 30.7 31.1 30.8 31.0 27.2
29.8 2.6
66.7 63.5 39.4 45.1 47.8 49.9 43.9 41.5 43.4 42.0 66.8 1.5
Pax7 33.6 34.2 26.4 28.0 24.0 40.0 40.0 29.1 32.7 34.9 23.1
31.4 8.4
43.1 41.5 59.5 55.8 65.0 39.8 39.8 53.3 45.1 40.1 67.0 4.4
Six4 26.8 27.2 25.2 26.3 26.4 29.7 27.2 28.0 26.4 28.8 24.4
26.9 2.6
51.2 48.3 61.8 54.5 53.4 31.0 48.5 42.5 54.0 36.9 67.9 1.4
Pitx2 40.0 40.0 28.7 25.1 25.2 34.5 40.0 24.1 34.5 32.5 20.8
31.4 10.6
40.3 40.3 52.6 59.7 59.5 41.4 40.3 61.6 41.2 45.2 67.9 5.1
Dlx3 32.6 35.9 31.9 31.0 30.2 36.0 34.2 32.9 30.0 33.8 28.2
32.4 4.2
48.5 37.0 51.7 55.8 59.6 37.0 40.9 46.9 60.8 42.6 69.2 2.1
Ehox-pending 33.0 35.3 31.2 30.0 29.2 33.1 40.0 29.3 31.9 32.1 26.9
32.0 5.1
43.8 35.5 51.2 56.1 59.8 43.5 35.5 59.3 48.2 47.7 69.2 2.4
Vdr 29.7 30.0 27.1 29.3 30.7 30.1 31.3 29.5 30.2 25.8 24.8
28.9 4.2
46.1 44.7 59.4 48.1 41.4 44.4 38.2 47.3 43.9 65.7 70.8 2.0
Barx1 34.7 40.0 40.0 32.6 40.0 40.0 40.0 34.9 40.0 30.1 29.7
36.5 6.9
45.6 44.0 44.0 56.1 44.0 44.0 44.0 44.7 44.0 68.8 71.1 2.0
Nr0b1 33.4 33.8 31.2 30.6 30.6 35.3 34.7 28.0 33.8 40.0 25.7
32.5 6.8
45.2 43.7 52.6 54.7 54.8 39.7 40.6 63.7 43.9 39.7 71.4 2.9
Msx1 31.1 30.7 29.5 29.0 30.1 29.7 28.1 29.5 26.5 29.1 25.1
29.0 3.8
37.3 39.7 47.0 49.6 43.4 45.3 55.0 46.7 64.4 49.0 72.6 1.7
Six2 27.9 28.6 29.2 31.6 29.9 33.7 31.7 31.9 31.6 27.9 23.4
29.8 6.4
56.9 54.4 52.2 43.4 49.4 35.3 42.8 42.2 43.0 56.8 73.7 2.7
Rax 33.2 34.7 40.0 32.5 35.2 37.2 34.2 27.2 37.1 34.3 24.9
33.7 8.8
48.9 44.4 43.4 51.0 43.4 43.4 45.7 67.0 43.4 45.5 73.8 3.3
Nr1h3 27.5 28.2 27.0 27.2 26.0 26.3 27.7 26.5 23.9 25.7 22.2
26.2 4.0
42.6 38.0 45.0 43.9 51.0 49.8 41.1 48.4 63.4 52.8 73.9 1.7
Gata2 26.1 26.5 24.7 25.6 25.2 26.8 26.4 26.8 26.0 26.5 23.3
25.8 2.6
47.0 43.4 60.6 52.6 55.8 40.2 44.5 40.1 48.1 43.0 74.8 1.0
Runx3 34.6 34.9 33.2 34.5 33.5 34.6 34.8 34.9 32.1 32.9 29.8
33.6 3.9
43.4 41.4 53.1 44.3 50.6 43.5 42.2 41.4 59.9 54.9 75.3 1.5
Crem 24.7 24.5 26.3 25.8 26.6 26.0 25.4 25.6 26.0 25.3 22.8
25.4 2.6
56.6 58.4 41.2 45.4 37.4 43.8 49.4 47.7 44.0 50.7 75.5 1.0
Cebpd 26.0 26.2 25.4 25.9 26.1 26.3 24.2 26.4 25.7 25.0 22.7
25.5 2.7
45.1 43.2 50.9 45.9 43.6 41.8 61.4 40.7 47.2 54.6 75.7 1.1
Six6 40.0 40.0 35.0 40.0 40.0 40.0 33.0 25.4 40.0 31.8 20.1
35.0 14.9
44.3 44.3 44.3 44.3 44.3 44.3 48.6 64.5 44.3 51.0 75.7 4.7
Esr1 28.4 28.7 27.4 28.3 28.2 30.5 29.2 28.2 30.9 30.3 23.7
28.5 4.8
50.7 48.8 56.0 51.0 51.9 39.5 46.5 51.9 37.3 40.2 76.0 1.9
Bcl3 31.5 33.4 30.4 31.1 31.4 31.7 33.1 30.2 30.7 31.2 26.4
31.0 4.6
47.1 36.3 53.5 49.4 47.9 46.3 37.8 54.5 51.9 48.7 76.4 1.7
Meox1 29.0 29.2 27.2 28.0 28.7 28.7 28.1 28.5 27.4 27.5 24.8
27.9 3.1
40.9 39.2 56.1 49.1 43.2 43.3 48.5 45.0 54.6 53.1 76.8 1.2
Nr5a1 34.4 40.0 34.8 39.0 40.0 35.4 34.1 31.9 40.0 34.7 29.6
35.8 6.2
48.0 44.2 45.6 44.2 44.2 44.2 49.6 63.0 44.2 45.8 76.9 1.6
Six1 29.0 30.0 29.3 31.2 29.1 33.2 30.9 31.8 28.2 29.1 22.2
29.4 7.3
51.6 47.9 50.7 43.6 51.4 36.2 44.5 41.1 54.7 51.4 76.9 2.7
Foxl2 31.1 29.5 30.0 30.5 30.5 31.8 31.1 31.8 30.6 31.3 27.0
30.5 3.5
45.1 57.8 53.4 49.6 49.9 39.6 45.5 39.5 49.1 43.7 76.9 1.3
Hesx1 34.1 34.8 33.2 33.6 32.4 40.0 40.0 35.0 33.4 32.6 28.5
34.3 5.8
46.0 42.1 51.1 48.8 55.8 41.2 41.2 41.5 50.3 54.8 77.1 1.8
Pitx1 34.9 36.2 33.4 32.2 28.4 36.2 35.6 35.1 36.5 33.1 21.8
33.0 11.3
44.1 44.0 48.1 51.1 60.6 44.0 44.0 44.0 44.0 48.7 77.6 3.9
Lhx3 32.5 32.2 28.2 30.7 31.8 33.8 32.5 31.4 30.1 32.4 22.0
30.7 8.7
44.3 45.1 58.0 49.9 46.3 39.9 44.1 47.8 51.9 44.6 78.1 3.1
Six5 29.8 30.8 29.6 29.2 29.5 30.2 29.8 30.2 28.8 29.4 25.9
29.4 3.4
46.6 38.0 48.1 51.6 49.1 43.3 46.2 43.5 55.2 50.0 78.4 1.2
Tbx19 35.2 33.7 35.3 35.6 40.0 40.0 34.8 29.4 40.0 34.4 22.7
34.6 11.9
45.0 48.6 45.0 45.0 45.0 45.0 45.7 60.2 45.0 46.6 78.6 3.7
Nr1h4 31.1 32.2 30.7 31.5 31.6 30.8 30.5 31.7 30.9 32.0 27.1
30.9 3.9
48.4 40.7 51.7 45.3 44.9 50.9 53.1 44.5 49.8 41.6 79.1 1.3
Irf6 28.6 29.7 29.2 29.3 29.6 29.4 29.5 30.7 28.2 30.0 23.9
28.9 5.0
51.7 45.3 48.5 47.9 46.0 47.0 46.8 39.4 54.0 43.8 79.4 1.7
Ascl1 24.7 24.9 25.4 25.2 25.8 25.7 24.7 26.2 25.4 25.8 21.0
25.0 3.9
51.8 50.6 46.5 48.2 43.8 44.7 52.3 41.0 46.9 44.2 79.8 1.3
Pit1 32.5 34.0 34.7 34.6 40.0 34.9 34.4 40.0 40.0 31.7 20.0
34.2 14.2
51.0 47.4 45.7 46.0 45.0 45.2 46.5 45.0 45.0 52.7 80.6 4.2
Prop1 40.0 40.0 40.0 35.2 40.0 40.0 40.0 40.0 40.0 40.0 28.8
38.5 9.7
46.8 46.8 46.8 46.8 46.8 46.8 46.8 46.8 46.8 46.8 81.6 1.8